Polystepha pilulae (Beutenmuller, 1892)    Synonyms: Oak Leaf Gall Midge.
Kingdom Animalia   >   Phylum Arthropoda   >   Class Insecta   >   Order Diptera   >   Family Cecidomyiidae   >   Genus Polystepha   

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The galls of the midge Polystepha pilulae in Baltimore Co., Maryland (8/31/2014). Photo by Richard Orr. (MBP list)

Polystepha pilulae in Dorchester Co., Maryland (9/14/2014). Verified by Charley Eiseman/BugGuide. Photo by Mark Etheridge. (MBP list)

Polystepha pilulae galls in Harford Co., Maryland (8/25/2019). Verified by Charley Eiseman/BugGuide. Photo by Dave Webb. (MBP list)

A Polystepha pilulae leaf gall measuring 3 mm in diameter in Harford Co., Maryland (6/9/2018). Photo by Dave Webb. (MBP list)

The galls of Polystepha pilulae on Southern Red Oak in Somerset Co., Maryland (9/15/2013). Photo by Jim Brighton. (MBP list)

The galls of Polystepha pilulae in Baltimore City, Maryland (9/22/2011). Determined by Charley Eiseman/BugGuide. Photo by Thomas Wilson. (MBP list)

The galls of Polystepha pilulae in Baltimore City, Maryland (9/22/2011). Determined by Charley Eiseman/BugGuide. Photo by Thomas Wilson. (MBP list)

Polystepha pilulae in Prince George's Co., Maryland (9/13/2013). (c) treichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Allegany Co., Maryland (8/23/2019). (c) Andy Gougherty, some rights reserved (CC BY-NC). Photo by Andy Gougherty via iNaturalist. (MBP list)

Polystepha pilulae in Howard Co., Maryland (9/28/2019). (c) Judy Gallagher, some rights reserved (CC BY-NC). Photo by Judy Gallagher. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (9/20/2015). (c) Carrie Seltzer, some rights reserved (CC BY-NC). Photo by Carrie Seltzer via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (9/20/2015). (c) Carrie Seltzer, some rights reserved (CC BY-NC). Photo by Carrie Seltzer via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (10/2/2017). (c) mmn noriko, some rights reserved (CC BY-NC). Photo by mmn_noriko via iNaturalist. (MBP list)

Polystepha pilulae in Howard Co., Maryland (9/28/2019). (c) Ashley M Bradford, some rights reserved (CC BY-NC). Photo by Ashley Bradford. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (6/26/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae galls in Harford Co., Maryland (8/13/2017). Photo by Dave Webb. (MBP list)

Polystepha pilulae in Washington Co., Maryland (7/7/2018). (c) Mark Etheridge, some rights reserved (CC BY-NC). Photo by Mark Etheridge. (MBP list)

Polystepha pilulae in Dorchester Co., Maryland (5/28/2021). (c) Mark Etheridge, some rights reserved (CC BY-NC). Photo by Mark Etheridge. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (10/2/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Washington Co., Maryland (7/7/2018). (c) Mark Etheridge, some rights reserved (CC BY-NC). Photo by Mark Etheridge. (MBP list)

Polystepha pilulae in Dorchester Co., Maryland (5/28/2021). (c) Mark Etheridge, some rights reserved (CC BY-NC). Photo by Mark Etheridge. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (10/2/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Dorchester Co., Maryland (5/28/2021). (c) Mark Etheridge, some rights reserved (CC BY-NC). Photo by Mark Etheridge. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (10/2/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (Date obscured). (c) Jennifer Wright, some rights reserved (CC BY-NC). Photo by Jennifer Wright via iNaturalist. (MBP list)

Polystepha pilulae in Dorchester Co., Maryland (9/4/2021). (c) Jim Brighton, some rights reserved (CC BY-NC). Photo by Jim Brighton. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (9/17/2017). (c) Elissa Totin, some rights reserved (CC BY). Photo by Elissa Totin via iNaturalist. (MBP list)

Polystepha pilulae in Dorchester Co., Maryland (8/19/2021). (c) Mark Etheridge, some rights reserved (CC BY-NC). Photo by Mark Etheridge. (MBP list)

A Polystepha pilulae larva in Harford Co., Maryland (10/14/2021). Verified by Yann Kemper/iNaturalist. Photo by Dave Webb. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (8/2/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (8/2/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (9/26/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (8/2/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (10/2/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (11/9/2021). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (6/23/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (7/6/2021). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (11/9/2021). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Harford Co., Maryland (8/16/2022). (c) kidneymoth, some rights reserved (CC BY-NC). Photo by kidneymoth via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (7/6/2021). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (11/9/2021). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/16/2021). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Prince George's Co., Maryland (7/16/2022). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Prince George's Co., Maryland (8/11/2022). (c) Benjamin Burgunder, some rights reserved (CC BY). Photo by Benjamin Burgunder via iNaturalist. (MBP list)

Polystepha pilulae in Garrett Co., Maryland (7/13/2023). (c) Mark Etheridge, some rights reserved (CC BY-NC). Photo by Mark Etheridge. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (6/13/2023). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Garrett Co., Maryland (Date obscured). (c) Brady O'Brien, some rights reserved (CC BY-NC). Photo by brady_obrien via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (7/6/2021). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (11/9/2021). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (7/6/2021). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (8/4/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (6/26/2022). (c) Wayne Longbottom, some rights reserved (CC BY-NC). Photo by Wayne Longbottom. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (8/17/2022). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Queen Anne's Co., Maryland (9/6/2021). (c) Jim Brighton, some rights reserved (CC BY-NC). Photo by Jim Brighton. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (8/4/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Queen Anne's Co., Maryland (9/6/2021). (c) Jim Brighton, some rights reserved (CC BY-NC). Photo by Jim Brighton. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (6/26/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Queen Anne's Co., Maryland (9/6/2021). (c) Jim Brighton, some rights reserved (CC BY-NC). Photo by Jim Brighton. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/20/2021). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Allegany Co., Maryland (6/19/2022). (c) Anne DeNovo, some rights reserved (CC BY-NC). Photo by Anne DeNovo via iNaturalist. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (8/17/2022). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Queen Anne's Co., Maryland (9/6/2021). (c) Jim Brighton, some rights reserved (CC BY-NC). Photo by Jim Brighton. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (11/9/2021). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (9/27/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Harford Co., Maryland (8/16/2022). (c) kidneymoth, some rights reserved (CC BY-NC). Photo by kidneymoth via iNaturalist. (MBP list)

Polystepha pilulae in Baltimore Co., Maryland (9/21/2022). (c) Zachary Wimberly, some rights reserved (CC BY). Photo by Zachary Wimberly via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (9/29/2022). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (11/9/2021). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/20/2021). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (6/24/2022). (c) Matthew Beziat, some rights reserved (CC BY-NC). Photo by Matthew Beziat. (MBP list)

Polystepha pilulae in Calvert Co., Maryland (10/4/2022). (c) Matthew Beziat, some rights reserved (CC BY-NC). Photo by Matthew Beziat. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (6/24/2022). (c) Matthew Beziat, some rights reserved (CC BY-NC). Photo by Matthew Beziat. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (9/29/2022). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/11/2021). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (8/4/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (10/12/2021). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (7/15/2021). (c) Matthew Beziat, some rights reserved (CC BY-NC). Photo by Matthew Beziat. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/11/2021). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Garrett Co., Maryland (9/3/2022). (c) maryland adventure babe, some rights reserved (CC BY-NC). Photo by maryland_adventure_babe via iNaturalist. (MBP list)

Polystepha pilulae in Worcester Co., Maryland (7/23/2022). (c) Jim Brighton, some rights reserved (CC BY-NC). Photo by Jim Brighton. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/11/2021). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Allegany Co., Maryland (6/19/2022). (c) Anne DeNovo, some rights reserved (CC BY-NC). Photo by Anne DeNovo via iNaturalist. (MBP list)

Polystepha pilulae in Worcester Co., Maryland (7/23/2022). (c) Jim Brighton, some rights reserved (CC BY-NC). Photo by Jim Brighton. (MBP list)

Polystepha pilulae in Garrett Co., Maryland (Date obscured). (c) Josh Emm, some rights reserved (CC BY-NC). Photo by Josh Emm. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (8/11/2022). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/24/2022). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/24/2022). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Prince George's Co., Maryland (7/30/2021). (c) Matthew Beziat, some rights reserved (CC BY-NC). Photo by Matthew Beziat. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (6/13/2023). (c) Katja Schulz, some rights reserved (CC BY). Photo by Katja Schulz via iNaturalist. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/24/2022). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (11/9/2021). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/20/2021). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (6/19/2022). (c) Matthew Beziat, some rights reserved (CC BY-NC). Photo by Matthew Beziat. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (9/29/2022). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Allegany Co., Maryland (6/19/2022). (c) Anne DeNovo, some rights reserved (CC BY-NC). Photo by Anne DeNovo via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (10/21/2016). (c) gail, some rights reserved (CC BY-NC). Photo by gail via iNaturalist. (MBP list)

Polystepha pilulae in Garrett Co., Maryland (Date obscured). (c) Josh Emm, some rights reserved (CC BY-NC). Photo by Josh Emm. (MBP list)

Polystepha pilulae in Prince George's Co., Maryland (10/21/2022). (c) Jenny Glenn, some rights reserved (CC BY-NC). Photo by Jenny Glenn via iNaturalist. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (8/17/2022). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/20/2021). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (9/29/2022). (c) Emily Summerbell, some rights reserved (CC BY). Photo by Emily Summerbell, Ph.D. via iNaturalist. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/24/2022). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/11/2021). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Anne Arundel Co., Maryland (9/16/2021). (c) Timothy Reichard, all rights reserved. Photo by Timothy Reichard. (MBP list)

Polystepha pilulae in Baltimore Co., Maryland (10/3/2022). (c) Ron Wertz, some rights reserved (CC BY-NC-SA). Photo by Ron Wertz via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (7/24/2023). (c) Annie Weissman, some rights reserved (CC BY). Photo by wearethechampignons via iNaturalist. (MBP list)

Polystepha pilulae in Montgomery Co., Maryland (7/24/2023). (c) Annie Weissman, some rights reserved (CC BY). Photo by wearethechampignons via iNaturalist. (MBP list)


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